It has been suggested that guttalatus and zonatus could be simply location/color variants of the same species, much in the same way melanurus and synspilla are.
Or it may be that the geographic separation of certain populations, is allowing the evolution or formation of a number of new species from a single ancestor.
Much in the same way that Herichthys carpintus, and Herichthys tamasopoensis, although until recently thought to be location variants of the same species, have now become separated into 2.
I guess it depends on whether you are a lumper, or splitter.
Beyond the taxonomic question, I have kept and spawned zonatus, and found them to be exceedingly beautiful, but one of the most aggressive Vieja/Paraneetroplus(?) with any other cichlid, For me, they seem to best be kept, in a very large, species only tank.
Herichthys tamasopoensis
Herichthys carpintus location "chairel"