I know you don't want to hear this, but I'm just trying to help, I don't want to see you waste your time and money. I build and repair boats for a living so keeping water at bay is what I do.
Just because it doesn't leak up to a certain point and leaks above a certain point doesn't mean the leak is between those two points and when you say that filling the tank more produces multiple leak all that says to me is that the cracks are opening up more and more and allowing more water out.
I think you are very naive to say there is absolutely no flex in the tank, no matter how heavily built a structure is the is always flex, especially in a wooden structure. Just the sheer weight of water is enough to compress the wood enough to open the seams up. You would be amazed how much you can compress softwoods.
Another possibility is voids within the plywood, all though I don't think you would see any evidence of these for quite some time.
I understand your frustration, but 'glassing is the only way to guarantee a water tight tank, you could have the whole thing done in a day, and it will last forever.
I wish you the best of luck in whatever course of action you decide to take.