I think Marine is a different ball game. Rivers and lakes are much different.If i take a net to the beach i end up with bucket loads of shrimp in less than an hour. If that were a ray it would gorgd itself possibly to death as some eels do.
I feed my rays lots especially if i see them getting skinny due to breeding attempts and try to alter food accordingly but tbey usually have bumps in which case i reduce feeding slightly just because the gas produced (bloated stomach) is probably uncomfortable.
I have a wild male 10+ yrs old and i expect he will have allsorts wrong if you do testes but hes not fat and imo could do with some weight on.